At the level of the outer kinetochore, one finds SUMOylated Ndc80 and NUF2 [35,36], as well as the kinesins CENPE/KIF10 [36] and KIF18A [47], the RNA binding protein NKAP [48], the Ran GTPase activating protein RanGAP1 [49], the ANAPC4 subunit of the APC/C [50,51], along with the protein kinases BUB1B/BubR1 [52] and MPS1/TTK [53].
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