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  • SUMO chains are able to recruit SUMO-targeted ubiquitin ligases (STUbLs) like RNF4 (Ring Finger Protein 4, Slx5/Slx8 heterodimer in S. cerevisiae) and RNF11/Arkadia, a class of ubiquitin E3 enzymes that ubiquitylate SUMOylated proteins [97].
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