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Within these ranges, seven different geographical units were considered for the study, six from the Eastern and Central Pyrenees (Catalonia, NE Spain): Freser-Setcases National Game Reserve (PyFS), Cadí National Game Reserve (PyC), Cerdanya-Alt Urgell National Game Reserve (PyCAU), Boumort National Game Reserve (PyB), Alt Pallars National Game Reserve (PyAP) and Vall Aran Game Reserve (PyVA); and one in the Eastern Cantabrian Mountains (León, N Spain), the Natural Protected Area of Picos de Europa (CmPE) (Fig 1).
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